For species with "black howler" in their common names, see Howler monkey.
Black howler[1]
Male
Female Both photographed at Pittsburgh Zoo
Conservation status
Near Threatened (IUCN 3.1)[2]
Scientific classification
Domain:
Eukaryota
Kingdom:
Animalia
Phylum:
Chordata
Class:
Mammalia
Order:
Primates
Suborder:
Haplorhini
Infraorder:
Simiiformes
Family:
Atelidae
Genus:
Alouatta
Species:
A. caraya
Binomial name
Alouatta caraya
(Humboldt, 1812)
Black Howler range
The black howler (Alouatta caraya) or black-and-gold howler,[2] is among the largest New World monkeys and a member of the Alouatta genus.[3][4] The black howler is distributed in areas of South America such as Paraguay, southern Brazil, eastern Bolivia, northern Argentina, and Uruguay.[5][6] This species is sexually dimorphic, with adult males having entirely black fur and adult females and babies of both sexes having an overall golden colouring; which emphasizes black-and-gold in the name.[6] The IUCN Red List has classed the black howler as Near Threatened as a result of a recent population reduction due to a variety of human-caused factors.[2]
Two females in a tree branch, using their strong prehensile tail as a safety fifth limb, in Pantanal, Bolivia
^Groves, C. P. (2005). Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore: Johns Hopkins University Press. p. 148. ISBN 0-801-88221-4. OCLC 62265494.
^ abcBicca-Marques, J.C.; Rumiz, D.I.; Ludwig, G.; Rímoli, J.; Martins, V.; da Cunha, R.G.T.; Alves, S.L.; Valle, R.R.; Miranda, J.M.D.; Jerusalinsky, L.; Messias, M.R.; Cornejo, F.M.; Boubli, J.P.; Cortes-Ortíz, L.; Wallace, R.B.; Talebi, M.; de Melo, F.R. (2021) [amended version of 2020 assessment]. "Alouatta caraya". IUCN Red List of Threatened Species. 2021: e.T41545A190414715. doi:10.2305/IUCN.UK.2021-1.RLTS.T41545A190414715.en. Retrieved 3 September 2022.
^Almeida, Marco Antônio Barreto de; Santos, Edmilson dos; Cardoso, Jader da Cruz; Fonseca, Daltro Fernandes da; Noll, Carlos Alberto; Silveira, Vivian Regina; Maeda, Adriana Yurika; Souza, Renato Pereira de; Kanamura, Cristina; Brasil, Roosecelis Araújo (1 January 2012). "Yellow fever outbreak affecting Alouatta populations in southern Brazil (Rio Grande do Sul State), 2008-2009". American Journal of Primatology. 74 (1): 68–76. doi:10.1002/ajp.21010. PMID 22020690. S2CID 20240047.
^Cortés-Ortiz, Liliana; Rylands, Anthony B.; Mittermeier, Russell A. (2015), Kowalewski, Martín M.; Garber, Paul A.; Cortés-Ortiz, Liliana; Urbani, Bernardo (eds.), "The Taxonomy of Howler Monkeys: Integrating Old and New Knowledge from Morphological and Genetic Studies", Howler Monkeys: Adaptive Radiation, Systematics, and Morphology, New York, NY: Springer, pp. 55–84, doi:10.1007/978-1-4939-1957-4_3, ISBN 978-1-4939-1957-4, retrieved 11 April 2022
^Alesci, Marco; Smith, Rebecca L.; Ayala Santacruz, Jorge Damian; Camperio Ciani, Andrea (1 March 2022). "Attitudes towards urban howler monkeys (Alouatta caraya) in Paraguay". Primates. 63 (2): 161–171. doi:10.1007/s10329-022-00975-5. ISSN 1610-7365. PMC 8898239. PMID 35142940.
^ abPérez‐Granados, Cristian; Schuchmann, Karl‐Ludwig (2021). "Passive acoustic monitoring of the diel and annual vocal behavior of the Black and Gold Howler Monkey". American Journal of Primatology. 83 (3): e23241. doi:10.1002/ajp.23241. ISSN 0275-2565. PMID 33539555. S2CID 231818221.
blackhowler (Alouatta caraya) or black-and-gold howler, is among the largest New World monkeys and a member of the Alouatta genus. The blackhowler is...
Howler monkeys (genus Alouatta, monotypic in subfamily Alouattinae) are the most widespread primate genus in the Neotropics and are among the largest of...
The brown howler (Alouatta guariba), also known as brown howler monkey, is a species of howler monkey, a type of New World monkey that lives in forests...
The mantled howler (Alouatta palliata) is a species of howler monkey, a type of New World monkey, from Central and South America. It is one of the monkey...
In Arkansas folklore, the Howler is a legendary creature said to dwell in the Ozarks.[citation needed] According to tradition, the creature is bear-like...
black fur, often with paler markings. These are arboreal and diurnal animals, with most species restricted to dense rain forest, although some howler...
of coyotes howling at night Problems playing this file? See media help. Howler monkey growls Problems playing this file? See media help. The long-distance...
the Coiba Island howler (Alouatta coibensis) is often recognized, but some authorities treat it as a subspecies of the mantled howler, (A. palliata). A...
The Colombian red howler or Venezuelan red howler (Alouatta seniculus) is a South American species of howler monkey, a type of New World monkey, found...
The Amazon blackhowler (Alouatta nigerrima) is a species of howler monkey, a type of New World monkey, endemic to the south-central Amazon in Brazil....
of the howler monkey genus, the Mexican howler is sympatric with the Guatemalan blackhowler, A. pigra, in Tabasco, Mexico. The Mexican howler differs...
Island howler (Alouatta coibensis) is a type of howler monkey, a type of New World monkey, endemic to Panama. Although the Coiba Island howler has been...
howlers, including adult males, while male harpy eagles tend to focus on juveniles. In one study, breeding harpy eagles hunted Yucatán blackhowler (Alouatta...
The ursine howler (Alouatta arctoidea) is a species of howler monkey native to Venezuela and possibly Colombia. It is sometimes considered a subspecies...
The Guyanan red howler (Alouatta macconnelli) is a species of howler monkey, a type of New World monkey, native to Suriname, Guyana, Trinidad, French Guiana...
Garber, Paul A. (2012). "Collective group movement and leadership in wild blackhowler monkeys (Alouatta pigra)". Behavioral Ecology and Sociobiology. 67: 31–41...
have a thick brown coat with dark gray appendages. The stomach area is black and heads are light brown. The fur color is the same for both males and...
acrobatically through the trees using their tails. Atelines, along with the related howler monkeys, are the largest of the New World monkeys. They live in groups,...
fruit, leaves, gum or insects. For example, leaf eaters such as howler monkeys, black-and-white colobuses and sportive lemurs have extended digestive...
shade of grey or black as well as brown, and are darker on the head and underparts. The face is naked of any fur and is typically black. Most male woolly...
can support its own weight. The body is long and slender and covered with black hair, which is coarse and often looks matted. Adult body length ranges between...
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