An African elephant in Tanzania, with visible tusks
Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with narwhals, chevrotains, musk deer, water deer, muntjac, pigs, peccaries, hippopotamuses and walruses, or, in the case of elephants, elongated incisors. Tusks share common features such as extra-oral position, growth pattern, composition and structure, and lack of contribution to ingestion. Tusks are thought to have adapted to the extra-oral environments, like dry or aquatic or arctic.[1] In most tusked species both the males and the females have tusks although the males' are larger. Most mammals with tusks have a pair of them growing out from either side of the mouth. Tusks are generally curved and have a smooth, continuous surface. The male narwhal's straight single helical tusk, which usually grows out from the left of the mouth, is an exception to the typical features of tusks described above. Continuous growth of tusks is enabled by formative tissues in the apical openings of the roots of the teeth.[2][3]
Other than mammals, dicynodonts are the only known vertebrates to have true tusks.[4]
^Nasoori, Alireza (2020). "Tusks, the extra-oral teeth". Archives of Oral Biology. 117: 104835. doi:10.1016/j.archoralbio.2020.104835. PMID 32668361. S2CID 220585014.
^"Tusk". The Oxford English Dictionary. 2010.
^Konjević, Dean; Kierdorf, Uwe; Manojlović, Luka; Severin, Krešimir; Janicki, Zdravko; Slavica, Alen; Reindl, Branimir; Pivac, Igor (4 April 2006). "The spectrum of tusk pathology in wild boar (Sus scrofa L.) from Croatia" (PDF). Veterinarski Arhiv. 76 (suppl.) (S91–S100). Retrieved 9 January 2011.
^Whitney, M. R.; Angielczyk, K. D.; Peecook, B. R.; Sidor, C. A. (2021). "The evolution of the synapsid tusk: Insights from dicynodont therapsid tusk histology". Proceedings of the Royal Society B: Biological Sciences. 288 (1961). doi:10.1098/rspb.2021.1670. PMC 8548784. PMID 34702071.
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