Ferns that grow with a trunk elevating the fronds above ground level
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A tree fern near Belles, DominicaAlsophila sp. tree ferns overlooking a valley in Misamis Oriental, Philippines
The tree ferns are arborescent (tree-like) ferns that grow with a trunk elevating the fronds above ground level, making them trees. Many extant tree ferns are members of the order Cyatheales, to which belong the families Cyatheaceae (scaly tree ferns), Dicksoniaceae, Metaxyaceae, and Cibotiaceae. It is estimated that Cyatheales originated in the early Jurassic,[1][2] and is the third group of ferns known to have given rise to tree-like forms. The others are the extinct Tempskya of uncertain position,[3] and Osmundales where the extinct Guaireaceae and some members of Osmundaceae also grew into trees. In addition there were the Psaroniaceae and Tietea in the Marattiales, which is the sister group to most living ferns including Cyatheales.
Other ferns which are also tree ferns, are Leptopteris and Todea in the family Osmundaceae, which can achieve short trunks under a metre tall. Fern species with short trunks in the genera Blechnum, Cystodium and Sadleria from the order Polypodiales, and smaller members of Cyatheales like Calochlaena, Cnemedaria, Culcita (mountains only tree fern), Lophosoria and Thyrsopteris are also considered tree ferns.
^Sosa, V.; Ornelas, J. F.; Ramírez-Barahona, S.; Gándara, E. (2016). "Historical reconstruction of climatic and elevation preferences and the evolution of cloud forest-adapted tree ferns in Mesoamerica". PeerJ. 4: e2696. doi:10.7717/peerj.2696. PMC 5119233. PMID 27896030.
^Sosa, Victoria; Ornelas, Juan Francisco; Ramírez-Barahona, Santiago; Gándara, Etelvina (2016). "Historical reconstruction of climatic and elevation preferences and the evolution of cloud forest-adapted tree ferns in Mesoamerica". PeerJ. 4. Chronogram of the Cyatheaceae and other tree fern lineages. doi:10.7717/peerj.2696. ISSN 2167-8359. PMC 5119233. PMID 27896030.
^Martínez, Leandro C.A.; Olivo, Mariana S. (August 2015). "Tempskya in the Valanginian of South America (Mulichinco Formation, Neuquén Basin, Argentina) — Systematics, palaeoclimatology and palaeoecology". Review of Palaeobotany and Palynology. 219: 116–131. Bibcode:2015RPaPa.219..116M. doi:10.1016/j.revpalbo.2015.04.002. hdl:11336/49538.
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