Various specimens of Notocupes from the Middle Jurassic of China and mid-Cretaceous Burmese amber
Scientific classification
Domain:
Eukaryota
Kingdom:
Animalia
Phylum:
Arthropoda
Class:
Insecta
Order:
Coleoptera
Suborder:
Archostemata
Family:
incertae sedis
Genus:
†Notocupes Ponomarenko, 1964
Synonyms
SinocupesLin, 1976
ChengdecupesHong, 1983
?ConexicoxaLin, 1986
EchinocupsKirejtshuk and Jarzembowski, 2020
AmblommaTan et al., 2005
OvatocupesTan and Ren, 2006
Notocupes is an extinct genus of medium-sized archostematan beetles from the Mesozoic Era of Eurasia, including over 50 described species. Historically, the genus was classified as a member of the family Ommatidae, but the presence of characters such as the horizontal mandibular cutting edge, separated procoxae and overlapping abdominal sternites indicate that the genus may have a closer affinity with the family Cupedidae.[1]Notocupes is considered to be a junior synonym of Zygadenia by Kirejtshuk (2020),[2] but other researchers suggest to reserve the genus Zygadenia as a form taxon for isolated elytra that probably belong to the genus Notocupes, while retaining Notocupes as a valid genus for complete body fossils.[3][4][5] Most species of Notocupes were described from compression fossils. An additional three species were described from Cenomanian-aged Burmese amber, which were treated as a separate genus, Echinocups, by Kirejtshuk (2020), but Li et al. (2023) consider Echinocups to be a junior synonym of Notocupes.[6]Notocupes has a flattened body, which may suggest that it occupied narrow habitats, such as living under bark. Some species had serrated/spined margins of the carapace, which may have served as a defense against predators, or served as camouflage to resemble bark.[1]
Strelnikova & Yan (2023) advise splitting up the genus Notocupes into four genera: Notocupes, Rhabdocupes, Conexicoxa and the new genus Brachilatus.[7]
^ abLi, Yan-Da; Tihelka, Erik; Yamamoto, Shûhei; Newton, Alfred F.; Xia, Fang-Yuan; Liu, Ye; Huang, Di-Ying; Cai, Chen-Yang (2023-08-22). "Mesozoic Notocupes revealed as the sister group of Cupedidae (Coleoptera: Archostemata)". Frontiers in Ecology and Evolution. 11. doi:10.3389/fevo.2023.1015627. ISSN 2296-701X.
^Kirejtshuk, Alexander G. (2020). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. Bibcode:2020Geosc..10...73K. doi:10.3390/geosciences10020073. ISSN 2076-3263.
^Strelnikova, O.D.; Yan, E.V. (2021). "Redescriptions of beetles of the Notocupes generic complex (Coleoptera: Archostemata: Ommatidae) from the Lower Cretaceous of Buryatia". Palaeoentomology. 4 (5): 499–514. doi:10.11646/palaeoentomology.4.5.15. S2CID 240527664.
^Strelnikova, O.D.; Yan, E.V. (2023). "Redescriptions of the Triassic Notocupes beetles (Archostemata: Ommatidae) from Kyrgyzstan and South Kazakhstan". Palaeoentomology. 6 (2): 174–190. doi:10.11646/palaeoentomology.6.2.9. S2CID 258403180.
^Soo Bin Lee; Gi Soo Nam; Yan-Da Li (2022). "A new species of Notocupes (Coleoptera: Archostemata) from the Lower Cretaceous (Albian) Jinju Formation in South Korea". Cretaceous Research. 140: 105357. Bibcode:2022CrRes.14005357L. doi:10.1016/j.cretres.2022.105357. S2CID 252147645.
^Li, Y.-D.; Tihelka, E.; Newton, A. F.; Huang, D.-Y.; Cai, C.-Y. (2023). "New species of Notocupes (Coleoptera: Archostemata) from the Middle Jurassic Daohugou beds, with discussion on the generic circumscription". Palaeoentomology. 6 (4): 398–415. doi:10.11646/palaeoentomology.6.4.11.
^Strelnikova, O.D.; Yan, E.V. (2023). "On splitting of the genus Notocupes (Coleoptera: Archostemata): new data on morphology and taxonomy". Far Eastern Entomologist. 488: 1–28. doi:10.25221/fee.488.1.
probably belong to the genus Notocupes, while retaining Notocupes as a valid genus for complete body fossils. Most species of Notocupes were described from compression...
junior synonym of Notocupes) †?Eurydictyon Ponomarenko, 1969 – Early Jurassic (Kyrgyzstan) (possibly closely related to Notocupes) †Jarzembowskiops Kirejtshuk...
It was created in 2020 to house three species originally assigned to Notocupes, E. denticollis, E. neli and E. ohmkuhnlei The genus name refers to the...
minor Ponomarenko, 1966 Rhabdocupes may be closely related to the genus Notocupes. Ommatidae Species List Archived 2012-10-23 at the Wayback Machine at...
triassicus Ponomarenko, 1966 Notocupoides may be closely related to the genus Notocupes. Ommatidae Species List Archived 2012-10-23 at the Wayback Machine at...
found in Kyrgyzstan. Eurydictyon may be closely related to the genus Notocupes. Ommatidae Species List Archived 2012-10-23 at the Wayback Machine at...
taxon for isolated elytra that probably belong to the genus Notocupes, while retaining Notocupes as a valid genus for complete body fossils. Eleven species...
Late Triassic †Taxopsis Kirejtshuk et al., 2016 – Baltic amber, Eocene Notocupes, traditionally considered a member of Ommatidae, has been suggested to...
Fang-Yuan; Liu, Ye; Huang, Di-Ying; Cai, Chen-Yang (2023-08-22). "Mesozoic Notocupes revealed as the sister group of Cupedidae (Coleoptera: Archostemata)"...
assigned to Allophalerus †Tetraphalerus laetus Lin, 1976 assigned to Notocupes †Tetraphalerus latus Tan et al. 2007 assigned to Allophalerus †Tetraphalerus...
S2CID 236585083. Lee, Soo Bin; Nam, Gi Soo; Li, Yan-Da (2022). "A new species of Notocupes (Coleoptera: Archostemata) from the Lower Cretaceous (Albian) Jinju Formation...
beetles, 12 species): Lithocupes (L. gigas, L. incertus, L. punctatus), Notocupes? (N. laticella, N. rostratus, N. tenuis), Notocupoides? (N. capitatus...
H.; Liu, Y.; Chen, R.; Kong, F. (2020). "A new species of the genus Notocupes from mid-Cretaceous Burmese amber (Coleoptera: Archostemata: Ommatidae)"...
theropods, is presented by Peñalver et al. (2023). Li et al. (2023) interpret Notocupes as a close relative of the family Cupedidae rather than an ommatid. Alleged...
on beetle diversity, is published by Zhao et al. (2021). Revision of Notocupes undatabdominus and N. ? multituberatus from the Lower Cretaceous of South...
S2CID 247143688. Lee, S. B.; Nam, G. S.; Li, Y.-D. (2022). "A new species of Notocupes (Coleoptera: Archostemata) from the Lower Cretaceous (Albian) Jinju Formation...
Alfred F. Newton; Di-Ying Huang; Chen-Yang Cai (2023). "New species of Notocupes (Coleoptera: Archostemata) from the Middle Jurassic Daohugou beds, with...
Alfred F. Newton; Di-Ying Huang; Chen-Yang Cai (2023). "New species of Notocupes (Coleoptera: Archostemata) from the Middle Jurassic Daohugou beds, with...
species of Nephus Notocupes lini Sp. nov Valid Ponomarenko et al. Early Jurassic China Originally described as a species of Notocupes; Kirejtshuk (2020)...
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