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The primary characters of the American film franchise of Planet of the Apes are a combination of humans and intelligent apes with both species acting as protagonist and antagonist across the series in three disparate timelines. The original film and characters were based on French author Pierre Boulle's 1963 novel Planet of the Apes. These characters from the original story were added to with each addition to the series.
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University of New York Press. pp. 139–152. ISBN 9781438430294. Keegan, Rebecca (11 August 2011). "'Rise ofthePlanetoftheApes': 21 nods to classic 'Apes'"...