Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga.[1][2] The Coptoclavidae lived from the Late Triassic to the Early Cretaceous.[3] Coptoclavidae is a member of the adephagan clade Dytiscoidea,[4][5][6] which contains other aquatic beetles. Suggested reasons for their extinction to include the rise of teleost fish, or competition with Gyrinidae and Dytiscidae, which possess defensive secretions and sucking channels in the mandibles of larvae, which coptoclavids likely lacked. It has been suggested that the genus Timarchopsis and the subfamily Timarchopsinae are only distantly related to other coptoclavids based on cladistic analysis, with Timarchopsis being more closely related to geadephagans like carabids and trachypachids instead.[7] Another study also suggested similarly for Coptoclavisca and possibly other coptoclaviscines.[8]
^Tree of Life Web Project. 1995. Liadytidae. Liadytes. Version 1 January 1995 (temporary). [1].01.01 in The Tree of Life Web Project, [2]
^Ivan Löbl & Aleš Smetana (May 2003). "1". Catalogue of Palaearctic Coleoptera: Archostemata, Myxophaga, Adephaga. Vol. 1. Apollo Books. pp. 6–8. ISBN 978-87-88757-73-6.
^Grimaldi, David and Michael S. Engel (16 May 2005). Evolution of the Insects. Cambridge University Press. ISBN 0-521-82149-5.
^Ponomarenko, A. G., Arnoldy, L., Jerikin, V. V., & Nikritin, L. M. (1992). Mesozoic Coleoptera. Suborder Adephaga, 1.
^Wang, B.; Ponomarenko, A. G.; Zhang, H. -Ch. (15 December 2009). "A new coptoclavid larva (Coleoptera: Adephaga: Dytiscoidea) from the Middle Jurassic of China, and its phylogenetic implication". Paleontological Journal. 43 (6): 652. Bibcode:2009PalJ...43..652W. doi:10.1134/S0031030109060082. ISSN 1555-6174. S2CID 83756102.
^Bo, Wang; Ponomarenko, A. G.; Haichun, Zhang (2010). "Middle Jurassic Coptoclavidae (Insecta: Coleoptera: Dytiscoidea) from China: a Good Example of Mosaic Evolution". Acta Geologica Sinica – English Edition. 84 (4): 680–687. Bibcode:2010AcGlS..84..680W. doi:10.1111/j.1755-6724.2010.00272.x. ISSN 1755-6724. S2CID 83806777.
^Beutel, Rolf G.; Wang, Bo; Tan, Jing-Jing; Ge, Si-Qin; Ren, Dong; Yang, Xing-Ke (April 2013). "On the phylogeny and evolution of Mesozoic and extant lineages of Adephaga (Coleoptera, Insecta)". Cladistics. 29 (2): 147–165. doi:10.1111/j.1096-0031.2012.00420.x. PMID 34814378. S2CID 85234655.
^Kirejtshuk, Alexander G.; Ansorge, Jörg (17 May 2022). "An extraordinarily preserved new genus and species of Trachypachidae (Coleoptera, Adephaga) from the Early Jurassic of Germany and a review of fossil trachypachid genera". Historical Biology. 35 (6): 958–975. doi:10.1080/08912963.2022.2071709. ISSN 0891-2963. S2CID 248869983.
Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga. The Coptoclavidae lived from the Late Triassic to the Early Cretaceous...
Dytiscidae) were less, with the most widespread being the species of Coptoclavidae, which preyed on aquatic fly larvae. A 2020 review of the palaeoecological...
A. J. Ross, and P. Davidson. 2017. Mesozoic Holcoptera (Coleoptera: Coptoclavidae) from England and the United States. Proceedings of the Geologists'...
Dytiscidae) were less, with the most widespread being the species of Coptoclavidae, which preyed on aquatic fly larvae. There are many fossils of beetles...
Ponomarenko, 1994 Family †Parahygrobiidae Ponomarenko, 1977 Family †Coptoclavidae Ponomarenko, 1961 Family †Liadytidae Ponomarenko, 1977 Family Meruidae...
Coptoclavella is an extinct genus of beetles in the family Coptoclavidae. There are about seven described species in Coptoclavella. These seven species...
Andrew J.; Davidson, Philip (2017). "Mesozoic Holcoptera (Coleoptera: Coptoclavidae) from England and the United States". Proceedings of the Geologists'...
Coptoclavinae is a beetle subfamily in the family Coptoclavidae. Zhang, HaiChun; Wang, Bo; Fang, Yan (2010-12-01). "Evolution of insect diversity in the...
sp Both nymph and adult fossils found A member of the extinct family Coptoclavidae. Similar to C. longipoda, but assigned to a yet undescribed species...
Hispanoclavinae is an extinct subfamily of ground and water beetles in the family Coptoclavidae. "The Paleobiology Database, Hispanoclavinae subfamily". paleobiodb...
Ponomarenko; A.S. Bashkuev (2013). "New beetle larvae (Coleoptera: Coptoclavidae, Caraboidea, Polyphaga) from the Upper Triassic of Germany" (PDF). Russian...
(2022). "A new species of the beetle genus Holcoptera (Coleoptera: Coptoclavidae) from the Upper Triassic of east Ukraine". Russian Entomological Journal...
J. Ross; Philip Davidson (2017). "Mesozoic Holcoptera (Coleoptera: Coptoclavidae) from England and the United States". Proceedings of the Geologists'...
Ponomarenko, Prokin & Bashkuev Middle to Late Triassic Germany A member of Coptoclavidae. Genus includes Stargelytron larissae and Stargelytron altus. Stegobium...
Ponomarenko in Ponomarenko et al. Late Jurassic Mongolia A member of Coptoclavidae, a species of Coptoclavella. Crassisyne Gen. et sp. nov Valid Yan, Wang...
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