Not to be confused with Bulbourethral gland or Bulb of penis.
Erect bulbus glandis (1) in a Labrador Retriever
The bulbus glandis (also called a bulb or knot) is an erectile tissue structure on the penis of canid mammals.[1][2][3][4][5][6] During mating, immediately before ejaculation the tissues swell up to lock (tie) the male's penis inside the female. The locking is completed by circular muscles just inside the female's vagina; this is called "the knot" tightening thus preventing the male from withdrawing. The circular muscles also contract intermittently, which has the effect of stimulating ejaculation of sperm, followed by prostatic fluid, as well as maintaining the swelling of the penis and therefore the tie, for some time. For domestic dogs the tie may last up to half an hour or more, though usually less.[7] When male canines are sexually excited, the bulbus glandis may swell up inside the penile sheath, even if the dog has been neutered.[8]
The bulbus glandis also occurs in the penises of some pinnipeds, including South American fur seals.[9]
^Howard E. Evans; Alexander de Lahunta (7 August 2013). Miller's Anatomy of the Dog - E-Book. Elsevier Health Sciences. ISBN 978-0-323-26623-9.
^Kim Long (1996). Wolves: A Wildlife Handbook. Big Earth Pub. ISBN 978-1-55566-158-8. Retrieved 10 December 2012.
^L. David Mech (16 May 2012). Wolf. Random House Digital, Inc. pp. 172–. ISBN 978-0-307-81913-0. Retrieved 10 December 2012.
^Marshall Cavendish Corporation (1 September 2010). Mammal Anatomy: An Illustrated Guide. Marshall Cavendish. pp. 252–. ISBN 978-0-7614-7882-9. Retrieved 10 December 2012.
^Debra A. Carlson (2008). Reproductive Biology of the Coyote (Canis Latrans): Integration of Behavior and Physiology. pp. 67–. ISBN 978-0-549-50315-6. Retrieved 9 February 2013.
^H. M. Menino (2008). Darwin's Fox and My Coyote. University of Virginia Press. pp. 101–. ISBN 978-0-8139-2675-9. Retrieved 9 February 2013.
^Susan Long (2006). Veterinary Genetics and Reproductive Physiology. Churchill Livingstone Elsevier. ISBN 978-0-7506-8877-2.
^"Boy Dog Bumps - The Mysterious Bulbus Glandis". Clarendon Animal Care. March 8, 2018. Retrieved June 17, 2020.
^Machado, Alex Sander D.; Bas, Le; Miglino, Maria Angélica; Leiser, Rudolf; Papa, Paula de Carvalho (1990-01-06). "Genital morphology of the male South American fur seal (Arctocephalus australis) and biological implications". Pesquisa Veterinária Brasileira. 32: 67–78. doi:10.1590/S0100-736X2012001300013. ISSN 0100-736X.
The bulbusglandis (also called a bulb or knot) is an erectile tissue structure on the penis of canid mammals. During mating, immediately before ejaculation...
two sections: Behind the lower, long part (pars longa glandis) lies the "knot" (Bulbusglandis) which expands only after penetrating the vagina and causes...
Vaginismus". Both men served on that respected journal's editorial board. Bulbusglandis Copulatory tie Priapism Taylor, Kräupl K. (October 20, 1979). "Penis...
and the collector massages the dog's erect penis near the base of the bulbusglandis using the thumb and index finger. The dog begins pelvic thrusting movements...
(trabeculae). Canine penises have a structure at the base called the bulbusglandis. During copulation, the spotted hyena inserts his penis through the...
possible that they form vaginal plugs or retain sperm after copulation. Bulbusglandis Leonard, Janet; Cordoba-Aguilar, Alex (2010-07-16). The Evolution of...
glans penis is smooth and consists of two parts called the bulbusglandis and pars longa glandis. The glans of a fossa's penis extends about halfway down...
needed] The canine penis contains a baculum and a structure called the bulbusglandis that expands during copulation, forming a copulatory tie that lasts...
dogs are coitally tied for around 5–30 minutes because of the male's bulbusglandis swelling and the female's constrictor vestibuli contracting; the male...
mate with the vixens for several days, often in burrows. The male's bulbusglandis enlarges during copulation, forming a copulatory tie which may last...
fibers. The clitoral frenulum or frenum (frenulum clitoridis and crus glandis clitoridis in Latin; the former meaning "little bridle") is a medial band...
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